Some of the most destructive forest invasions happen entirely underground, in northern forests that spent more than 10,000 years with almost no earthworms. European nightcrawlers strip the duff layer northern hardwoods depend on, and Asian jumping worms, which breed without a mate and spread in mulch and bait, leave dry castings that won't hold water. Native wildflowers and ground-nesting birds fall, and disturbed soil opens the door to garlic mustard and buckthorn.
Whether the dramatic garden-soil transformation by jumping worms adds up to stand-scale forest collapse is unsettled; the Wisconsin arboretum has reported limited effects so far in its forest plots.